Thanks everyone! Looks like more agitation possibly with a brush might be a good place to start. And yes it is uneven development in mostly even toned skies, not underdevelopment.
I've attached a crop of a jpeg. The sky is clear blue, cloudless, with a very little haze near the horizon. Please note that the "spots" in the left hand sky are a result of the scanner platen and are NOT present on the negative.
Frank, the artifacts are visible in the negative.
Tony one-by-one from presoak to developer. Agitation is by fore-aft, side-to-side, and a "circular" or "rotational" rocking of the tray - I usually do some of all three every agitation cycle.
Sandy thanks for the info. My contrast is about right so I'll have to play with agitation methods.
And thanks everyone - this is just the kind of information needed to help solve problems.
Cheers, Steve
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